Experiment / E9L4QY9OPTargeted / Cap-STARR-seq

DENND1A-locus allele-specific targeted STARR-seq in H295R cells

Gene regulatory activity associated with polycystic ovary syndrome revealed DENND1A-dependent testosterone production

Genomic DNA from five healthy female 1000 Genomes individuals was enriched across the DENND1A locus, cloned into an episomal STARR-seq vector, pooled, and assayed in H295R cells. The processed table retains the 62 of 623 assayed variants with BIRD posterior probability greater than 0.90 for an allele-specific regulatory effect.

Processed tables are specific to each experiment. Column names, units, measurements, and table structure are not standardized across the database. Check this experiment’s column definitions and quality-control notes before comparing or combining data.

Perturbation & assay details

Basal / Untreated

A targeted, episomal STARR-seq adaptation. Genomes from three European-ancestry and two Han Chinese-ancestry healthy female 1000 Genomes individuals were sheared to approximately 200 bp, captured with Agilent SureSelect custom probes spanning GRCh38 chr9:123279654-124030107, cloned into the STARR-seq ORI vector, pooled equimolarly, and transfected into H295R cells (70 million cells and 140 micrograms per replicate; three biological replicates). Input and reporter libraries were sequenced, allele counts were obtained after WASP/Bowtie2 processing and samtools mpileup, and BIRD estimated theta as the alternate-allele transcriptional rate relative to the reference allele.

Processed data

50 rows per page. Click a cell to inspect its full value.

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Filters apply to this table only. The CSV download contains the complete processed table; filtered rows are available through the API.

Column dictionary · 20 definitions
variant_id
dbSNP rsID of the tested variant from Supplementary Data 11.
chromosome
GRCh38 chromosome for the tested variant, when supplied by Supplementary Data 11 or the matching Data 12 annotation.
position_hg38
1-based GRCh38 variant position; blank when the source did not supply a coordinate.
ref_allele
Reference allele reported in the source BIRD input/output summary.
alt_allele
Alternate allele reported in the source BIRD input/output summary.
alt_allele_frequency
Alternate-allele frequency (af) in the pooled 1000 Genomes source population.
minor_allele
Whether the minor allele is the reference allele (REF) or alternate allele (ALT), as reported by the source.
minor_allele_frequency
Minor-allele frequency (MAF) reported in Supplementary Data 11.
theta_alt_over_ref
BIRD posterior-median effect size theta, interpreted as alternate-allele transcriptional rate divided by reference-allele transcriptional rate.
theta_lower_95ci
Lower bound of the BIRD 95% credible interval for theta.
theta_upper_95ci
Upper bound of the BIRD 95% credible interval for theta.
posterior_probability
BIRD posterior probability P of a regulatory effect; rows were retained at P > 0.90.
standard_error
BIRD standard error for theta reported in Supplementary Data 11.
alternate_allele_activity
Derived direction relative to the alternate allele: increased for theta > 1, decreased for theta < 1, and unchanged for theta = 1.
minor_allele_activity
Derived direction for the minor allele, reversing the alternate-allele direction when the minor allele is REF.
orthogonal_overlap
ENCODE ATAC/DNase or enhancer-like element annotation joined from Supplementary Data 12, when available.
eqtl_gene
GTEx eQTL gene annotation joined from Supplementary Data 12, when available.
coordinate_source
Supplementary Data 11 when the source supplied coordinates; Supplementary Data 12 when its matching position filled missing Data 11 coordinates.
supplementary_annotation_status
Whether a matching rsID was present in Supplementary Data 12; Data 12-only variants were not added to the BIRD result set.
qc_status
Package retention status; PASS_PAPER_BIRD_P_GT_0.90 indicates the paper-defined posterior-probability filter and value/ID checks passed.

Quality control

The paper retained reads with MAPQ >= 30 outside centromeres and blacklist regions, used WASP and Bowtie2 for allele-aware alignment, marked duplicates with Picard, and assigned variant allele counts with samtools mpileup. BIRD was run with its standard model and a regulatory-effect threshold of 1.2; the source table contains 623 assayed variants. Package QC retained the paper-defined 62 variants with posterior probability P > 0.90, finite theta/credible interval/standard-error values, and unique variant IDs. Three retained variants lack genomic coordinates and allele fields in Supplementary Data 11; those missing source values remain blank rather than being imputed, except that the matching Supplementary Data 12 position was used for rs894139170.

Curation notes

This is the study's genuine allele-specific MPRA/STARR-seq experiment. The 62 retained rows are the paper's BIRD-positive variants from Supplementary Data 11, not the nearby PCOS GWAS association summary statistics. The raw_data folder includes the three input and three reporter RPKM tracks, the complete source workbook, and the five 1000 Genomes sample IDs. The alternate_allele_activity and minor_allele_activity fields are deterministic interpretations of BIRD theta; no new statistical effect estimates were recomputed from the RPKM tracks. Supplementary Data 11 and Data 12 are not perfectly identical: 59 high-probability rsIDs join directly, three Data 11 candidates have no Data 12 annotation, and three Data 12 annotations are not in the retained Data 11 P > 0.90 set.

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